Publications

This page contains a citation list of publications that have been authored by The Orianne Society staff.  Staff members are displayed in bold on each citation.  PDFs (click on the title to access) and links to publications are provided.

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Peer-Reviewed Articles​
Notes & County Records​
Reports
Other Publications

Peer-Reviewed Articles

84. Brooks, G. C., S. J. Goodman, W. M. Moore, H. C. Chandler, N. M. Caruso, J. A. Smith, T. A. Gorman, and C. A. Haas. 2025. Gopher Tortoise (Gopherus polyphemus) burrow associate communities are depauperate in modified landscapes. Community Ecology 26:223–237. https://doi.org/10.1007/s42974-025-00232-x

Orianne Society staff member checks camera traps at a drift fence. South Georgia.

83. Samuels, L.R.N., T. Wilcox, M. Hoffman, M. Elmore, R. Aldredge, B.S. Stegenga, J. Bogan Jr., M.A. Davis, S. Hertz, M.K. Schwartz, and H.C. Chandler. 2025. Comparison of camera traps, eDNA, and visual encounter surveys for threatened species detection. Journal of Nature Conservation 86:126948. https://doi.org/10.1016/j.jnc.2025.126948

Close up of an invasive pentastome, 1mm line for size reference.

82. Palmisano, J. N., M. Brennan, A. M. Durso, J. H. Kesselring, T. Morgan, Z. Lepera, B. S. Stegenga, E. E. Luckenbaugh, S. Tsoukias, S. L. McKnight, K. Titterington, L. Milligan, H. D. S. Walden, R. J. Ossiboff, A. E. Savage, and T. M. Farrell. 2025. Rapid spread of the invasive pentastome Raillietiella orientalis (Hett, 1915) in 14 new Florida counties and in pet trade snakes. BioInvasions Records 14:261–269. https://doi.org/10.3391/bir.2025.14.1.20

81. Samuels, L. R. N., H. C. Chandler, M. Hoffman, J. A. Kronenberger, M. Elmore, R. Aldredge, B. S. Stegenga, J. E. Bogan Jr., M. A. Davis, S. Hertz, M. K. Schwartz, and T. Wilcox. 2025. Persistence of reptile DNA in a terrestrial substrate: A case study using the Eastern Indigo Snake. Environmental DNA 7:e70053. https://doi.org/10.1002/edn3.70053

80. Chandler, H.C., N.M. Caruso, G.C. Brooks, and C.A. Haas. 2024. Wetland hydrology, not altered phenology, challenges Reticulated Flatwoods Salamander (Ambystoma bishopi) management under future climate change. Ichthyology & Herpetology 112:531–543. https://doi.org/10.1643/h2023052

79. Brooks, G.C., H.C. Chandler, C.A. Haas, and H.K. Kindsvater. 2024. Ten principles from evolutionary ecology for the effective conservation of reptiles and amphibians. Journal of Herpetology 58:1–20.  https://doi.org/10.1670/2330062

78. Chandler, H.C., B.S. Stegenga, Z.A. Cava, A. Colton, M.T. Holden, D. Hutto, Jr., T.M. Wantman, and J.D. Mays. 2024. Ghosts of the swamp: Challenges associated with surveying for Spotted Turtles (Clemmys guttata) in the southeastern United States. Northeastern Naturalist 31:C1–C16.  https://doi.org/10.1656/045.031.s1201

77. Chandler, H. C., D. L. McLaughlin, and C. A. Haas. 2024. Informing the conservation of ephemerally flooded wetlands using hydrologic regime and LiDAR-based habitat assessments. Wetlands 44:33. https://doi.org/10.1007/s13157-023-01767-3

76. Brown, G., J.D. Mays, H.C. Chandler, B.S. Stegenga, B. Kreiser, and D.J. Stevenson. 2023. Spotted Turtle (Clemmys guttata) Population Genetics in the Southeastern United States. Chelonian Conservation and Biology 22:127–136. https://doi.org/10.2744/CCB-1560.1

75. Chandler, H.C., and G.C. Brooks. 2023. The relationship between female body size and clutch size in flatwoods salamanders. Southeastern Naturalist 22:588–594. https://doi.org/10.1656/058.022.0411

74. Chandler, H.C., D. Steen, J. Blue, J.E. Bogan, M.R. Bolt, T. Brady, D.R. Breininger, J. Buening, M. Elliott, J. Godwin, C. Guyer, R.L. Hill, M. Hoffman, N.L. Hyslop, C.L. Jenkins, C. Lechowicz, M. Moore, R.A. Moulis, S. Piccolomini, R. Redmond, F.H. Snow, B.S. Stegenga, D.J. Stevenson, J. Stiles, S. Stiles, M. Wallace, J. Waters, M. Wines, and J.M. Bauder. 2023. Evaluating growth rates of captive, wild, and reintroduced populations of the imperiled Eastern Indigo Snake (Drymarchon couperi). Herpetologica 79:220–230. https://doi.org/10.1655/Herpetologica-D-22-00041

73. Brooks G.C., H.C. Chandler, Y. Jiao, D.C. Childs, and C.A. Haas. 2023. Predicting the population viability of an endangered amphibian under environmental and demographic uncertainty. Population Ecology. https://doi.org/10.1002/1438-390X.12172

72. Brooks, G.C., N.M. Caruso, H.C. Chandler, and C.A. Haas. 2023. Niche partitioning and the storage effect facilitate coexistence in an amphibian community. Ecology and Evolution 13:e10629. https://doi.org/10.1002/ece3.10629

71. Brooks, G.C., T.A. Gorman, K.C. Jones, H.C. Chandler, B.K. Rincon, M.A. Sission, J. Himes, and C.A. Haas. 2023. Removing duff layers in fire-suppressed wetlands can aid habitat restoration efforts. Wetlands 43:95. https://doi.org/10.1007/s13157-023-01739-7

70. Chandler, H.C., N.M. Caruso, D.L. McLaughlin, Y. Jiao, and C.A. Haas. 2023. Forecasting the flooding dynamics of flatwoods salamander breeding wetlands under future climate change scenarios. PeerJ 11:e16050. https://doi.org/10.7717/peerj.16050

69. Bolt, M.R., J.M. Bauder, M.L. Legare, C.L. Jenkins, B.B. Rothermel, and D.R. Breininger. 2023. Eastern Indigo Snake (Drymarchon couperi) shelter site use in peninsular Florida, USA, and implications for habitat conservation. Herpetological Conservation and Biology 18:362-373. https://www.herpconbio.org/Volume_18/Issue_2/Bolt_etal_2023.pdf

68. Stegenga, B.S., D.S. Stevenson, and H.C. Chandler. 2023. Observations of the Suwannee Alligator Snapping Turtle from the Okefenokee Swamp. Southeastern Naturalist 22:78-83. https://doi.org/10.1656/058.022.0sp1204

67. Roberts, H.P., L.L. Willey, M.T. Jones, D.I. King, T.S.B. Akre, J. Kleopfer, D.J. Brown, S.W. Buchanan, H.C. Chandler, P. deMaynadier, M. Winters, L. Erb, K.D. Gipe, G. Johnson, K. Lauer, E.B. Liebgold, J.D. Mays, J.R. Meck, J. Megyesy, J.L. Mota, N.H. Nazdrowicz, K.J. Oxenrider, M. Parren, T.S. Ransom, L. Rohrbaugh, S. Smith, D. Yorks, and B. Zarate. 2023. Effects of landscape structure and land use on turtle communities across the eastern United States. Biological Conservation 283: 110088. https://doi.org/10.1016/j.biocon.2023.110088

66. O’Hanlon, B., J.E. Bogan, J.C. Godwin, M. Hoffman, L.L. Smith, H.C. Chandler, and B.L. Talley. 2023. Cryptosporidium serpentis surveillance in free-ranging snakes to inform a reintroduction strategy for the Eastern Indigo Snake (Drymarchon couperi). Journal of Wildlife Diseases 59:176–180. https://doi.org/10.7589/JWD-D-22-00055

65. Roberts, H. P., L. L. Willey, M. T. Jones, T. S. B. Akre, D. I. King, J. Kleopfer, D. J. Brown, S. W. Buchanan, H. C. Chandler, P. G. deMaynadier, M. Winters, L. Erb, K. D. Gipe, G. Johnson, K. Lauer, E. B. Liebgold, J. D. Mays, J. R. Meck, J. Megyesy, J. L. Mota, N. H. Nazdrowicz, K. J. Oxenrider, M. Parren, T. S. Ransom, L. Rohrbaugh, S. Smith, D. Yorks, and B. Zarate. 2023. Is the future female for turtles? Climate change and wetland configuration predict sex ratios of a freshwater species. Global Change Biology 29:2643–2654. https://doi.org/10.1111/gcb.16625

64.  Bauder, J. M., H. C. Chandler, M. L. Elmore, and C. L. Jenkins. 2022. Incorporating habitat suitability, landscape distance, and resistant kernels to estimate conservation units for an imperiled terrestrial snake. Landscape Ecology 37:2519­–2533. https://doi.org/10.1007/s10980-022-01510-z

63.  Chandler, H.C., B.S. Stegenga, and J.D. Mays. 2022. Compensating for small body size: The reproductive ecology of southern Spotted Turtle (Clemmys guttata) populations. Ichthyology & Herpetology 110:268–277. https://doi.org/10.1643/h2021084

62.  Chandler, H. C., C. L. Jenkins, and J. M. Bauder. 2022. Accounting for geographic variation in species-habitat associations during habitat suitability modeling. Ecological Applications 32:e2504. https://doi.org/10.1002/eap.2504

61.  Enge, K. M., D. J. Stevenson, T. M. Thomas, G. R. Johnston, J. B. Jensen, B. S. Stegenga, H. C. Chandler, and P. E. Moler. 2021. Distribution and relative abundance of the Suwannee Alligator Snapping Turtle (Macrochelys suwanniensis). Chelonian Conservation and Biology 20:184–199. https://doi.org/10.2744/CCB-1488.1

60.  Lentz, T. B., M. C. Allender, S. Y. Thi, A. S. Duncan, A. X. Miranda, J. C. Beane, D. S. Dombrowski, B. R. Forester, C. K. Akcali, N. A. Shepard, J. E. Corey III, A. L. Braswell, L. A. Williams, C. R. Lawson, C. Jenkins, J. H. K. Pechmann, J. Blake, M. Hooper, K. Freitas, A. B. Somers, B. L. Stuart. 2021. Prevalence of Ranavirus, Batrachochytrium dendrobatidis, B. salamandrivorans, and Ophidiomyces ophiodiicola in amphibians and reptiles of North Carolina, USA. Herpetological Review 52:285–293.
59.  Snow, F.H., A.B. Safer, D.J. Stevenson, and H.C. Chandler. 2021. Use of a novel refuge type by Eastern Indigo Snakes (Drymarchon couperi) in Georgia. Southeastern Naturalist 20:308–314. https://doi.org/10.1656/058.020.0208
58.  Chandler, H. C., J. C. Colón-Gaud, T. A. Gorman, K. Carson, and C. A. Haas. 2021. Does long-term fire suppression impact leaf litter breakdown and aquatic invertebrate colonization in pine flatwoods wetlands? PeerJ 9:e12534. https://doi.org/10.7717/peerj.12534
57.  Bauder, J. M., W. E. Peterman, S. F. Spear, C. L. Jenkins, A. R. Whiteley, and K. McGarigal. 2021. Multiscale assessment of functional connectivity: Landscape genetics of eastern indigo snakes in an anthropogenically fragmented landscape in central Florida. Molecular Ecology 30:3422–3438. https://doi.org/10.1111/mec.15979
56.  Bauder, J. M., D. R. Breininger, M. R. Bolt, M. L. Legare, C. L. Jenkins, B. B. Rothermel, and K. McGarigal. 2020. Movement barriers, habitat heterogeneity or both? Testing hypothesized effects of landscape features on home range sizes in Eastern Indigo Snakes. Journal of Zoology 311:204–216. https://doi.org/10.1111/jzo.12777
55.  Chandler, H. C., S. J. Goodman, J. A. Smith, T. A. Gorman, and C. A. Haas. 2020. Understanding occupancy patterns in a low-density Gopher Tortoise (Gopherus polyphemus) population. Herpetological Conservation and Biology 15:536–546. http://www.herpconbio.org/Volume_15/Issue_3/Chandler_etal_2020.pdf
54.  Chandler, H. C., B. S. Stegenga, and D. J. Stevenson. 2020. Thermal ecology of Spotted Turtles (Clemmys guttata) in two southern populations. Copeia 108:737–745. https://doi.org/10.1643/CE-19-315
53.  Haynes, E., H. C. Chandler, B. S. Stegenga, L. Adamovicz, E. Ospina, D. Zerpa-Catanho, D. J. Stevenson, and M. C. Allender. 2020. Ophidiomycosis surveillance of snakes in Georgia, USA reveals new host species and taxonomic associations with disease. Scientific Reports 10:10870. https://doi.org/10.1038/s41598-020-67800-1
52. Deitloff, J., E. Myers, S. Spear, D. Stevenson, and C. Guyer. 2019. Multiple paternity and heritability of color in Drymarchon couperi (Eastern Indigo Snakes). Herpetologica 75:224–232. https://doi.org/10.1655/D-18-00027

51. Chandler, H. C., B. S. Stegenga, and D. J. Stevenson. 2019. Movement and space use in southern populations of Spotted Turtles (Clemmys guttata). Southeastern Naturalist 18:602–618. https://doi.org/10.1656/058.018.0411

50.  Brooks, G. C., J. A. Smith, E. A. Frimpong, T. A. Gorman, H. C. Chandler, and C. A. Haas. 2019. Indirect connectivity estimates of amphibian breeding wetlands from spatially explicit occupancy models. Aquatic Conservation Marine and Freshwater Ecosystems 29:1815–1825. https://doi.org/10.1002/aqc.3190.
49.  Chandler, H. C., M. C. Allender, B. S. Stegenga, E. Haynes, E. Ospina, and D. J. Stevenson. 2019. Ophidiomycosis prevalence in Georgia’s Eastern Indigo Snake (Drymarchon couperi) populations. PLoS ONE 14: e0218351. https://doi.org/10.1371/journal.pone.0218351
48.  Folt, B., J. Bauder, S. Spear, D. Stevenson, M. Hoffman, J. R. Oaks, P. L. Wood Jr., C. Jenkins, D. A. Steen, and C. Guyer. 2019. Taxonomic and conservation implications of population genetic admixture, mito-nuclear discordance, and male-biased dispersal of a large endangered snake, Drymarchon couperi. PLoS ONE 14:e0214439. https://doi.org/10.1371/journal.pone.0214439
47.  Bauder, J. M., D. R. Breininger, M. R. Bolt, M. L. Legare, C. L. Jenkins, B. B. Rothermel, and K. McGarigal. 2018. Multi-level, multi-scale habitat selection by a wide-ranging, federally threatened snake. Landscape Ecology 33:743–763. https://doi.org/10.1007/s10980-018-0692-2
45.  Chandler, H. C., D. L. McLaughlin, T. A. Gorman, K. J. McGuire, J. B. Feaga, and C. A. Haas. 2017. Drying rates of ephemeral wetlands: Implications for breeding amphibians. Wetlands 37:545–557. https://doi.org/10.1007/s13157-017-0889-1
44.  Chandler, H. C., D. J. Stevenson, J. D. Mays, B. S. Stegenga, W. H. Vaigneur, and M. D. Moore. 2017. A new trap design for catching small Emydid and Kinosternid turtles. Herpetological Review 16:261–282.
43.  Stevenson, D. J., and H. C. Chandler. 2017. The herpetofauna of conservation lands along the Altamaha River, Georgia. Southeastern Naturalist 16:261–282. https://doi.org/10.1656/058.016.0216
42. Maritz, B., J. Penner, M. Martins, J. Crnobrnja-Isailović, S. Spear, L. R.V. Alencar, J. Sigala-Rodriguez, K. Messenger, R. W. Clark, P. Soorae, L. Luiselli, C. Jenkins, and H. W. Greene. 2016. Identifying global priorities for the conservation of vipers. Biological Conservation 204:94-102. http://doi.org/10.1016/j.biocon.2016.05.004
41. Goldberg, C., C. R. Turner, K. Deiner, K. E.Klymus, P. F. Thomsen, M. A. Murphy, S. F. Spear, A. McKee, S. J. Oyler-McCance, R. S. Cornman, M. B. Laramie, A. R. Mahon, R. F. Lance, D. S. Pilliod, K. M. Strickler, L. P. Waits, A. K. Fremier, T. Takahara, J. E. Herder, and P. Taberlet. 2016. Critical considerations for the application of environmental DNA methods to detect aquatic species. Methods in Ecology and Evolution 7:1299–1307. https://doi.org/10.1111/2041-210X.12595

40. Pierson, T. W., A. M. McKee, S. F. Spear, J. C. Maerz, C. D. Camp, and T. C. Glenn. 2016. Detection of an enigmatic plethodontid salamander using environmental DNA. Copeia 2016:78–82. http://doi.org/10.1643/CH-14-202

39.  Bauder, J. M., D. R. Breininger, M. R. Bolt, M. L. Legare, C. L. Jenkins, B. B. Rothermel, and K. McGarigal. 2016. Seasonal variation in Eastern Indigo Snake (Drymarchon couperi) movement patterns and space use in Peninsular Florida at multiple temporal scales. Herpetologica 72:214–226. https://doi.org/10.1655/Herpetologica-D-15-00039.1
38.  Bauder, J. M., D. R. Breininger, M. R. Bolt, M. L. Legare, C. L. Jenkins, B. B. Rothermel, and K. McGarigal. 2016. The influence of sex and season on conspecific spatial overlap in a large, actively-foraging colubrid snake. PLoS ONE 11:e0160033. https://doi.org/10.1371/journal.pone.0160033
37.  Richardson, J. L., S. P. Brady, I. J. Wang, and S. F. Spear. 2016. Navigating the pitfalls and promise of landscape genetics. Molecular Ecology 24:849–863. https://doi.org/10.1111/mec.13527
36.  Knafo, S. E., T. M. Norton, M. Mitchell, D. J. Stevenson, N. Hyslop, R. Poppenga, et al. 2016. Health and nutritional assessment of free‐ranging Eastern Indigo Snakes (Drymarchon couperi) in Georgia, United States. Journal of Zoo and Wildlife Medicine 47:1000–1012. https://doi.org/10.1638/2015-0127.1

35. McKee, A. M., S. F. Spear, and T. W. Pierson. 2015. The effect of dilution and the use of a post-extraction nucleic acid purification column on the accuracy, precision, and inhibition of environmental DNA samples. Biological Conservation 183:70–76. http://doi.org/10.1016/j.biocon.2014.11.031

34. McKee, A. M., D. L. Calhoun, W. J. Barichivich, S. F. Spear, C. S. Goldberg, and T. C. Glenn. 2015. Assessment of environmental DNA for detecting presence of imperiled aquatic amphibian species in isolated wetlands. Journal of Fish and Wildlife Management 6:498–510. https://doi.org/10.3996/042014-JFWM-034

33. Steen, D. A., M. Barbour, C. J. W. McClure, K. P. Wray, J. N. Macey, and D. J. Stevenson. 2015. Landscape scale habitat selection of Harlequin Coralsnakes (Micrurus fulvius) in three large, protected areas in the southeastern United States. Copeia 103:1037–1042. https://doi.org/10.1643/CE-15-235

32.  Dierenfeld, E. S., T. M. Norton, N. L. Hyslop, and D. J. Stevenson. 2015. Nutrient composition of prey items consumed by free-ranging Drymarchon couperi (Eastern Indigo Snakes). Southeastern Naturalist 14:551560. https://doi.org/10.1656/058.014.0311

31.  Stevenson, D. J., J. B. Jensen, E. A. Schlimm, and M. Moore. 2015. The distribution, habitat use, activity, and status of the Spotted Turtle (Clemmys guttata) in Georgia. Chelonian Conservation and Biology 14:136–142. https://doi.org/10.2744/CCB-1130.1

30. Bauder, J. M., D. R. Breininnger, M. R. Bolt, M. L. Legare, C. L. Jenkins, and K McGarrigal. 2015. The role of the bandwidth matrix in influencing kernel home range estimates for snakes using VHF telemetry data. Wildlife Research 42:437–453. https://doi.org/10.1071/WR14233
29. Spear, S. F., S. A. Cushman, and B. H. McRae. 2015. Resistance surface modeling in landscape genetics. Pages 129-148 in N. Balkenhol, S. A. Cushman, A. T. Storfer, and L. P. Waits, editors. Landscape Genetics: Concepts, Methods, Applications. John Wiley & Sons, Ltd. Chichester, United Kingdom. https://doi.org/10.1002/9781118525258.ch08
27.  Bauder, J. M., J. N. Macey, K. M. Stohlgren, A. Day, F. Snow, A. Safer, R. Redmond, J. M. Waters, M. Wallace, Sr., and D. J. Stevenson. 2015. Factors influencing the display of multiple defensive behaviors in Eastern Indigo Snakes (Drymarchon couperi). Herpetological Conservation and Biology 10:559–571. http://www.herpconbio.org/Volume_10/Issue_2/Bauder_etal_2015.pdf

26. Steen, D. A., C. J. W. McClure, J. C. Brock, D. C. Rudolph, J. B. Pierce, J. R. Lee, W. J. Humphries, B. B. Gregory, W. B. Sutton, L. L. Smith, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2014. Snake co-occurrence patterns are best explained by habitat and hypothesized effects of interspecific interactions. Journal of Animal Ecology 83:286–295. https://doi.org/10.1111/1365-2656.12121

25. Steen, D. A., C. J. W. McClure, W. B. Sutton, D. C. Rudolph, J. B. Pierce, J. R. Lee, L. L. Smith, B. B. Gregory, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2014. Copperheads are common when kingsnakes are not: Relationships between the abundances of a predator and one of their prey. Herpetologica 70:69–76. https://doi.org/10.1655/HERPETOLOGICA-D-13-00064

24.  Bauder, J. M., C. Castellano, J. B. Jensen, D. J. Stevenson, and C. L. Jenkins. 2014. Comparison of movements, body weight, and habitat selection between translocated and resident gopher tortoises. The Journal of Wildlife Management 78:1444–1455. https://doi.org/10.1002/jwmg.790

22.  Hyslop, N. L., J. M. Meyers, R. J. Cooper, and D. J. Stevenson. 2014. Effects of body size and sex of Drymarchon couperi (Eastern Indigo Snake) on habitat use, movements, and home range size in Georgia. Journal of Wildlife Management 78:101–111. https://doi-org.ezproxy.lib.vt.edu/10.1002/jwmg.645

19.  Stiles, S., J. C. Stiles, J. C. Godwin, C. Jenkins, E. M. Rush, B. Lock, V. M. Johnson, M. Wines, and C. Guyer. 2013. Repatriation of eastern indigo snakes to conservation lands in South Alabama, USA. Pages 37–41 in P. S. Soorae, editor. Global Re-introduction Perspectives: 2013. IUCN/SSC Re-introduction Specialist Group (RSG), Gland, Switzerland.
18.  Steen, D. A., D. J. Stevenson, J. C. Beane, J. D. Willson, M. J. Aresco, J. C. Godwin, S. P. Graham, L. L. Smith, J. M. Howze, D. C. Rudolph, J. B. Pierce, J. R. Lee, B. B. Gregory, J. Jensen, S. H. Stiles, J. A. Stiles, N. H. Nazdrowicz, and C. Guyer. 2013. Terrestrial Movement of the Red-bellied Mudsnake (Farancia abacura) and Rainbow Snake (F. erytrogramma). Herpetological Review 44:208–213.
17.  Enge, K. M., D. J. Stevenson, M. J. Elliott, and J. M. Bauder. 2013. The historical and current distribution of the Eastern Indigo Snake (Drymarchon couperi). Herpetological Conservation and Biology 8:288–307. http://www.herpconbio.org/Volume_8/Issue_2/Enge_etal_2013.pdf

16. Stevenson, D. J., and J. Stackhouse. 2012. The amphibians and reptiles of the Altamaha River, Georgia. Reptiles & Amphibians 19:170–186. https://doi.org/10.17161/randa.v19i3.13900

15. Stevenson, D. J., G. Greer, and M. J. Elliott. 2012. The distribution and habitat of Centruroides hentzi (Banks) (Scorpiones, Buthidae) in Georgia. Southeastern Naturalist 11:589–598. https://doi.org/10.1656/058.011.0404

13.  Hyslop, N. L., D. J. Stevenson, J. N. Macey, L. D. Carlile, C. L. Jenkins, J. A. Hostetler, and M. K. Oli. 2012. Survival and population growth of a long-lived threatened snake species, Drymarchon couperi (Eastern Indigo Snake). Population Ecology 54:145–156. https://doi.org/10.1007/s10144-011-0292-3
12.  Graves, T.A., T. N. Wasserman, M. C. Ribeiro, E. L. Landguth, S. F. Spear, N. Balkenhol, C. B. Higgins, M. J. Fortin, S. A. Cushman, and L. P. Waits. 2012. The influence of landscape characteristics and home-range size on the quantification of landscape-genetics relationships. Landscape Ecology 27:253–266. https://doi.org/10.1007/s10980-011-9701-4
11.  Steen, D. A., C. J. W. McClure. J. C. Brock, D. C. Rudolph, J. B. Pierce, J. R. Lee, W. J. Humphries, B. B. Gegory, W. B. Sutton, L. L. Smith, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2012. Landscape-level influences of terrestrial snake occupancy within the southeastern United States. Ecological Applications 22:1084–1097. https://doi.org/10.1890/11-1777.1
10.  Shamblin, B. M., T. I. Alstad, D. J. Stevenson, J. N. Macey, F. H. Snow, and C. J. Nairn. 2011. Isolation and characterization of microsatellite markers from the threatened Eastern Indigo Snake (Drymarchon couperi). Conservation Genetic Resources 3:303–306. https://doi.org/10.1007/s12686-010-9348-5

9. Anderson, C. D., B .K. Epperson, M. Fortin, R. Holderegger, P. M. A. James, M. S. Rosenberg, K. T. Scribner, and S. Spear. 2010. Considering spatial and temporal scale in landscape-genetic studies of gene flow. Molecular Ecology 19:3565–3575. https://doi.org/10.1111/j.1365-294X.2010.04757.x

8.  Storfer, A., M. A. Murphy, S. F. Spear, R. Holderegger, and L. P. Waits. 2010. Landscape genetics: Where are we now? Molecular Ecology 19:3496–3514. https://doi.org/10.1111/j.1365-294X.2010.04691.x
 
7.  Stevenson, D. J., K. R. Ravenscroft, R. T. Zappalorti, M. D. Ravenscroft, S. W. Weigley, and C. L. Jenkins. 2010. Using a wildlife detector dog for locating Eastern Indigo Snakes (Drymarchon couperi). Herpetological Review 41:437–442.

6.  Stevenson, D. J., M. R. Bolt, D. J. Smith, K. M. Enge, N. L. Hyslop, T. M. Norton, K. J. Dyer. 2010. Prey Records for the Eastern Indigo Snake (Drymarchon couperi). Southeastern Naturalist 9:1–18. https://doi.org/10.1656/058.009.0101.

5.  Spear, S. F., N. Balkenhol, M. Fortin, B. H. McRae, and K. Scribner. 2010. Use of resistance surfaces for landscape genetic studies: Considerations for parameterization and analysis. Molecular Ecology 19:3576–3591. https://doi.org/10.1111/j.1365-294X.2010.04657.x

4. Hyslop, N. L., J. M. Meyers, R. J. Cooper, and D. J. Stevenson. 2009. Indigo snake capture methods: Effectiveness of two survey techniques for Drymarchon couperi in Georgia. Florida Scientist 72:93–100.

2.  Stevenson, D. J., G. Beaton, and M. J. Elliott. 2009. Distribution, status and ecology of Cordulegaster sayi Selys in Georgia, USA (Odonata:Cordulegastridae). Bulletin of American Odonatology 11:20–25.
1.  Stevenson, D. J., K. M. Enge, L. D. Carlile, K. J. Dyer, T. M. Norton, N. L. Hyslop, and R. A. Kiltie. 2009. An Eastern Indigo Snake (Drymarchon couperi) mark-recapture study in southeastern Georgia. Herpetological Conservation and Biology 4:30–42. http://www.herpconbio.org/Volume_4/Issue_1/Stevenson_etal_2009.pdf?t=1528329600056

Natural History Notes & County Records

38.  Stegenga, B. S., Fedler, M. T., Stevenson, D. J., Landberg, T., and Palis, J. G. 2021. Pseudobranchus striatus (Northern Dwarf Siren). Reproduction. Herpetological Review 52:599-601.
37.  Stevenson, D.J., B.S. Stegenga, B. Rice, J. Barrett, J. Bolton, Z. Cava, H.C. Chandler, K.M. Stohlgren, T. Brock, C.D. Camp, J.B. Jensen, M.D. Moore, S. McGuire, C. Howe, F. Snow, J. Oguni, C. Coppola, and L.L. Smith. 2021. New County Records for amphibians and reptiles in Georgia, USA. Herpetological Review 52:350–359.
36.  Stevenson, D.J., B.S. Stegenga, H.C. Chandler, and R.D. Birkhead. 2021. Drymarchon couperi (Eastern Indigo Snake) – Distribution records. Herpetological Review 52:347.

35. Stevenson, D. J., G. Brown, H. Chandler, D. D. Dye II, C. Garza, M. McWhorter, M. Moore, and A. Thomas. 2018. Recent noteworthy distribution records for Deinopis spinosa (Marx, 1889) (Araneae: Deinopidae) in the Southeastern United States. Southeastern Naturalist 17:N28-N33. https://doi.org/10.1656/058.017.0209

34.  Stevenson, D. J., J. D. Mays, and H. C. Chandler. 2018. Clemmys guttata (Spotted Turtle). Coloration. Herpetological Review 49:317–318.
33.  Chandler, H. C., B. S. Stegenga, D. J. Stevenson, and J. Y. Lamb. 2018. Desmognathus cf. auriculatus (Southern Dusky Salamander). First County Record. Herpetological Review 49:68.

32. Hammerstein, G., D. A. Steen, and D. J. Stevenson. 2015. Crotalus horridus (Timber Rattlesnake). Diet. Herpetological Review 46:640–641.

31.  Stevenson, D. J., C. L. Jenkins, K. M. Stohlgren, J. B. Jensen, D. L. Bechler, I. Deery, D. duff, S. P. Graham, R. Herrington, P. Higgins, R. V. Horan III, C. Kelehear, K. Dylan, K. Kincaid, L. D. Mcbrayer, M. Moore, C. Muise, T. Muise, A. Muise, J. Oguni, E. M. Schlimm, and W. Vaigneur. 2015. Significant new records of amphibians and reptiles from Georgia, USA. Herpetological Review 46:597–601.

30. Warfel, T., M. Cage, and D. J. Stevenson. 2015. Micrurus distans distans (West Mexican Coralsnake). Diet. Herpetological Review 46:275.

28. Stevenson, D. J. 2014. Eastern Coachwhip (Coluber flagellum flagellum) preys on hatchling Gopher Tortoises (Gopherus polyphemus). Bulletin of the Chicago Herpetological Society 49:93–94.

27. Jenkins, C. L., B. Hudson, K. M. Stohlgren, S. F. Spear, D. J. Stevenson, and C. D. Camp. 2014. Desmognathus folkertsi (Dwarf Black-bellied Salamander. Predation. Herpetological Review 45:472–473.

26. Stevenson, D. J., J. B. Jensen, P. G. Frank Jr., W. Bond Jr., D. Duff, M. Moore, K. Nelson, J. Oguni, R. Preston Jr., M. Riddle, F. Snow, and K. M. Stohlgren. 2014. New county records for amphibians and reptiles of Georgia, USA. Herpetological Review 45:102–104.

25. Horan III, R. V., C. Rentz, and D. J. Stevenson. 2013. Drymarchon couperi (Eastern Indigo Snake). First County Record. Herpetological Review 44:109.

24. Stevenson, D. J. 2013. Drymarchon couperi (Eastern Indigo Snake). First County Record. Herpetological Review 44:109.

23. Stevenson, D. J. 2013. Bachman’s Sparrow use of a Gopher Tortoise burrow as an escape refuge. The Oriole 78:13–15.

22. Parker, J.M., S. F. Spear, S. Oyler-McCance. 2012. Crotalus oreganus concolor (midget faded rattlesnake) nursery aggregation. Herpetological Review 43:658–659.

21.Jensen, J. B., and D. J. Stevenson. 2012. Plestiodon egregius (Mole Skink). Mating activity. Herpetological Review 43:650.

20.  Stevenson, D. J. 2012. Farancia abacura (Mudsnake). First County Record. Herpetological Review 43:620.
 
19.  Stevenson, D. J., and P. G. Frank Jr. 2012. Farancia erytrogramma erytrogramma (Common Rainbow Snake). First County Record. Herpetological Review 43:620.

18. Day, A. J., and D. J. Stevenson. 2012. Hyla avivoca (Bird-voiced Treefrog). First County Record. Herpetological Review 3:613.

17. Butler, H., C. D. Camp, G. C. Greer, J. B. Jensen, D. Olive, D. J. Stevenson, and K. M. Stohlgren. 2012. New county records of amphibians and reptiles in Georgia, USA. Herpetological Review 43:453–456.

16. Stevenson, D. J., J. Parker, and M. P. Wallace. 2012. Pseudemys concinna concinna (Eastern River Cooter). New County Records. Herpetological Review 43:442–443.

15. Stevenson, D. J., and D. A. Steen. 2012. Nerodia erythrogaster erythrogaster (Red-bellied Watersnake). Necrophagy by dung beetles (Scarabaeinae). Herpetological Review 43:347.

14.  Bauder, J. M., J. N. Macey, M. P. Wallace, F. Snow, A. B. Safer, and D. J. Stevenson. 2012. Drymarchon couperi (Eastern Indigo Snake). Juvenile observations. Herpetological Review 43:343.

13. Parker, J., T. Warfel, M. Ishimatsu, and D. J. Stevenson. 2012. Storeria dekayi (Brownsnake). First County Record. Herpetological Review 43:107–108.

12. Stohlgren, K. M., and D. J. Stevenson. 2012. Stereochilus marginatus (Many-lined Salamander). First County Record. Herpetological Review 43:96.

11.  Beane, J. C., T. Pusser, F. Snow, K. V. Briggs, and D. J. Stevenson. 2011. Pseudotriton montanus (Mud Salamander). First County Record. Herpetological Review 42:384.

10. Carlile, L. D., T. A. Beaty, G. C. Hart, R. A. Brooks, and D. J. Stevenson. 2011. Heterodon platirhinos (Eastern Hog-nosed Snake). Maximum Size. Herpetological Review 42:291–292.

9. Stevenson, D. J., K. M. Stohlgren, E. M. Schlimm, J. M. Bauder, M. P. Wallace Sr., and W. Taylor. 2011. Additional geographic distribution records for amphibians and reptiles from Georgia, USA. Herpetological Review 42:255–257.

8. Jensen, J. B., D. J. Stevenson, S. Graham, M. P. Wallace Sr., C. Muise, T. Muise, A. B. Safer, F. Snow, R. Preston Jr., G. C. Greer, B. Hudson, and C. D. Camp. 2011. Additional distributional records of amphibians and reptiles in Georgia, USA. Herpetological Review 42:250–255.

7. Kellner, W. S., K. V. Briggs, J. M. Bauder, and D. J. Stevenson. 2011. Drymarchon couperi (Eastern Indigo Snake). First County Record. Herpetological Review 42:114.

6. Stevenson, D. J. 2010. Pituophis melanoleucus mugitus (Florida Pinesnake). First County Record. Herpetological Review 41:111.

5.  Stevenson, D. J., and N. L. Hyslop. 2010. Drymarchon couperi (Eastern Indigo Snake): Long-distance interpopulation movement. Herpetological Review 41:91–92.
 

4.  Stevenson, D. J. 2010. Drymarchon couperi (Eastern Indigo Snake): Death feigning. Herpetological Review 41:92–93. https://doi.org/10.1656/058.009.0114

 3.  Powell, C., D. J. Stevenson, M. Smith, and J. B. Jensen. 2010. A new clutch size record for the Mud Snake (Farancia abacura). Southeastern Naturalist 9:177–178. https://doi.org/10.1656/058.009.0114.

2. Wallace, M. P., D. J. Stevenson, and L. V. LaClaire. 2009. Captive longevity and size records for the Peninsula Newt (Notophthalmus viridescens piaropicola) and Striped Newt (Notophthalmus perstriatus). Bulletin of the Chicago Herpetological Society 44:138–139.

1. Delaney, D. K., L. L. Pater, L. D. Carlile, D. J. Stevenson, and A. D. Walde. 2008. Red-cockaded Woodpecker (Picoides borealis) response to nest depredation by an Eastern Rat Snake (Elaphe alleghaniensis). Southeastern Naturalist 7:753–759. https://doi.org/10.1656/1528-7092-7.4.756

Reports

 
7.  Stevenson, D. J., E. M. Schlimm, K. M. Stohlgren, J. Hepinstall-Cymerman. 2014. 2013–2014 Georgia status surveys for the Florida Pine Snake (Pituophis melanoleucus mugitus) and the Southern Hognose Snake (Heterodon simus). 14 pages.
6.  Antonio, F., and R. A. Odum. 2012. Eastern Indigo Snake (Drymarchon couperi) population management plan 2010–2012. Association of Zoos and Aquariums. 15 pages.
4.  Spear, S. F., J. M. Parker, C. R. Peterson, and C. L. Jenkins. 2011. Conservation and management of Midget Faded Rattlesnakes. State Wildlife Grant Final Report. 37 pages.
3.  Spear, S. F. 2011. Habitat connectivity for western rattlesnake (Crotalus oreganus) in the Columbia Plateau ecoregion. Appendix A.9 Washington Connected Landscapes Project: Analysis of the Columbia Plateau Ecoregion. 27 pages.
2.  AZA Snake TAG. 2011. Eastern Indigo Snake (Drymarchon couperi) care manual. Association of Zoos and Aquariums, Silver Spring, Maryland. 57 pages.
1.  Antonio, F. 2010. Eastern Indigo Snake studbook (Drymarchon couperi), 2nd edition. Association of Zoos and Aquariums. 59 pages.

Other Publications by The Orianne Society Staff

 
20.  Chandler, H. C., T. A. Gorman, and C. A. Haas. 2016. The effects of crayfish predation and vegetation cover on tadpole growth, survival, and nonlethal injury. Journal of Herpetology 50:271–277. https://doi.org/10.1670/14-176
19.  Erwin, K. J., H. C. Chandler, J. G. Palis, T. A. Gorman, and C. A. Haas. 2016. Herpetofaunal communities in ephemeral wetlands embedded within longleaf pine flatwoods of the Gulf Coastal Plain. Southeastern Naturalist 15:431–447. https://doi.org/10.1656/058.015.0305
15.  Anderson, C. D., B. K. Epperson, M. Fortin, R. Holderegger, P. M. A. James, M. S. Rosenberg, K. T. Scribner, and S. Spear. 2010. Considering spatial and temporal scale in landscape-genetic studies of gene flow. Molecular Ecology 19:3565–3575. https://doi.org/10.1111/j.1365-294X.2010.04757.x
13.  Storfer, A., J. M. Eastman, and S. F. Spear. 2009. Modern molecular methods for amphibian conservation. Bioscience 59:559–571. https://doi.org/10.1525/bio.2009.59.7.7
12.  Rothermel, B. B., S. C. Walls, J. C. Mitchell, C. K. Dodd Jr., L. K. Irwin, D. E. Green, V. M. Vazquez, J. W. Petranka, and D. J. Stevenson. 2008. Widespread occurrence of the amphibian chytrid fungus Batrachochytrium dendrobatidis in the southeastern USA. Diseases of Aquatic Organisms 82:3–18. https://doi.org/10.3354/dao01974
10.  Jenkins, C. L., and C. R. Peterson. 2008. A trophic-based approach to the conservation biology of rattlesnakes: Linking landscape disturbance to rattlesnake populations. Pages 265–274 in W. K. Hayes, K. R. Beaman, M. D. Cardwell, and S. P. Bush, editors. The Biology of Rattlesnakes. Loma Linda University Press, Loma Linda, California.
8.  Storfer, A., M. A. Murphy, J. S. Evans, C. S. Goldberg, S. Robinson, S. F. Spear, R. Dezzani, E. Delmelle, L. Vierling, and L. P. Waits. 2007. Putting the ‘landscape’ in landscape genetics. Heredity 98:128–142. https://doi.org/10.1038/sj.hdy.6800917
4.  Spear, S. F., C. R. Peterson, M. D. Matocq, and A. Storfer. 2005. Landscape genetics of the blotched tiger salamander (Ambystoma tigrinum melanostictum). Molecular Ecology 14:2553–2564. https://doi.org/10.1111/j.1365-294X.2005.02573.x
3.  Stevenson, D. J., K. J. Dyer, B. A. Willis-Stevenson. 2003. Survey and monitoring of the Eastern Indigo Snake in Georgia. Southeastern Naturalist 2:393–408. https://doi.org/10.1656/1528-7092(2003)002[0393:SAMOTE]2.0.CO;2
1.  Jenkins, C. L., K. McGarigal, and L. R. Gamble. 2002. A comparison of aquatic surveying techniques used to sample Ambystoma opacum larvae. Herpetological Review 33:33–15.